* Channing Tatum as Conrad Hauser / Duke: The lead soldier. Lorenzo di Bonaventura wanted to cast Mark Wahlberg in the role when the script was not about the origin story, while the studio met with Sam Worthington when it was rewritten by Beattie in its final incarnation. Tatum had played a soldier in Stop-Loss, an anti-war film, and originally wanted no part in G.I. Joe, which he felt glorified war. Once he read the script though, he realized the franchise was a fantasy akin to X-Men, Mission: Impossible and Star Wars rather than a war film.
* Dennis Quaid as General Clayton Abernathy / Hawk: The team leader. Quaid described Hawk as "a cross between Chuck Yeager and Sgt. Rock and maybe a naïve Hugh Hefner". Quaid's son convinced him to take on the part, and the filmmakers enjoyed working with him so much that Stuart Beattie wrote "ten to fifteen more scenes" for the character. He filmed all his scenes within the first two months of production. Quaid is signed on for two sequels.
* Rachel Nichols as Shana M. O'Hara / Scarlett: She graduated college at age twelve and became the team's intelligence expert. Having left school so early, she does not understand men's attraction to her. Nichols was the first choice for the role. Nichols had dyed her blonde hair red – Scarlett's hair color – for her role in Star Trek, which she filmed before G.I. Joe. She burned herself filming an action sequence with Miller.
* Ray Park as Snake-Eyes: A mute and mysterious ninja commando. Like his character, Park is a martial arts expert and specifically practiced wushu for the role, as well as studying the character's comic book poses. Park had known of Snake-Eyes having played with the toys as a child, but he knew very little of the surrounding saga of G.I. Joe vs. Cobra, so he read the comics to further understand the character. He was nervous about wearing the mask, so he requested to practice wearing it at home. He found the full costume, including the visor, very heavy to wear and akin to a rubber band; he had to put effort into moving in it.
* Marlon Wayans as Wallace Weems / Ripcord: He has a crush on Scarlett, which she is oblivious to, and he does not realize she is in a relationship with Snake-Eyes. A fan of the franchise, Wayans was cast on the strength of his performance in Requiem for a Dream. Bonaventura said that film showed Wayans could be serious as well as funny.
* Adewale Akinnuoye-Agbaje as Hershel Dalton / Heavy Duty: An ordnance expert. Common was offered the role of Heavy Duty's cousin Roadblock, although Bonaventura previously indicated Heavy Duty was being used in that character's stead. Stuart Beattie ultimately chose to have Heavy Duty instead of Roadblock.
* Saïd Taghmaoui as Abel Shaz / Breaker: He is the team's communications specialist and hacker.
* Karolína Kurková as Courtney A. Kreiger / Cover Girl: Hawk's aide-de-camp.
* Brendan Fraser as a yet-unnamed motorcycle-riding Sergeant. It was initially rumored he would play Gung-Ho, but those rumors turned out false. According to Fraser himself, he plays a motorcycle-riding sergeant who is the descendant of Rick O'Connell, Fraser's own character from Sommers' The Mummy film series.
COBRA
* Christopher Eccleston as James McCullen Destro XXIV / Destro: A weapons designer and founder of the Military Armament Research Syndicate (MARS) and the main villain for the early part of the film. Irish actor David Murray was cast as Destro, but was forced to drop it when he had problems with his visa. Murray was later cast as James McCullen I in a flashback scene.
* Joseph Gordon-Levitt as The Doctor / Cobra Commander: USA Today reported Cobra Commander is a multiple role. Levitt wore a mask – which was redesigned from the comics because the crew found it too reminiscent of the Ku Klux Klan – and prosthetic makeup underneath of it. Upon seeing concept art of the role he was being offered, Levitt signed on because; "I was like, 'I get to be that? You're going to make that [makeup] in real life and stick it on me? Cool. Let me do it.' That's a once-in-lifetime opportunity." Levitt is a friend of Tatum and they co-starred in Stop-Loss and Havoc. His casting provided extra incentive for Tatum to join the film. Levitt described his vocal performance as being half reminiscent of Chris Latta's voice for the 1980s cartoon, but also half his own ideas, because he felt rendering it fully would sound ridiculous.
* Sienna Miller as Baroness Anastasia DeCobray / The Baroness: A spy. Years before the film, the Baroness was going to marry Duke, but he left her at the altar. Miller auditioned for the part because it did not involve "having a breakdown or addicted to heroin or dying at the end, something that was just maybe really great fun and that people went to see and actually just had a great time seeing". Miller prepared with four months of weight training, boxing sessions and learned to fire live ammunition, gaining five pounds of muscle. She sprained her wrist after slipping on a rubber bullet while filming a fight between the Baroness and Scarlett.
* Lee Byung-hun as Thomas Arashikage / Storm Shadow: Snake-Eyes' rival, both were close members of the Arashikage ninja clan. Lee said he did not know G.I. Joe because it is an unknown series in Korea. Sommers and Bonaventura told him not to watch any of the cartoons to prepare for the role. Lee was attracted to Storm Shadow's "dual personality", which he stated has "huge pride and honor".
* Arnold Vosloo as Zartan: An expert in makeup and disguises serving Destro.
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